Fullmetal Alchemist: Brotherhood (Japanese: 鋼の錬金術師 FULLMETAL ALCHEMIST, Hepburn: Hagane no Renkinjutsushi: Furumetaru Arukemisuto) is an anime series adapted from the Fullmetal Alchemist manga by Hiromu Arakawa. Produced by Bones, the series is directed by Yasuhiro Irie and written by Hiroshi Ōnogi. Fullmetal Alchemist: Brotherhood is the second anime television series based on Fullmetal Alchemist, the first being 2003's Fullmetal Alchemist. Unlike the previous adaptation, Brotherhood is an almost 1:1 adaptation directly following the original events of the manga. It was first announced in the manga series' 20th tankōbon volume. In Japan, it is differentiated from the 2003 series by the inclusion of the English language title. The series premiered on April 5, 2009, on MBS-TBS' Sunday 5:00 PM JST anime time block, replacing Mobile Suit Gundam 00, and ran weekly until airing its final episode on July 4, 2010. Voice actresses Romi Park and Rie Kugimiya reprised their roles as main characters Edward and Alphonse Elric, respectively.